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Which railroad service/network does Tesla use?

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They may well use several. Depends on where they are shipping. For instance, the UP (Union Pacific) doesn't run very much east past Chicago. So, if they are sending cars from the west coast to the east coast, those cars will likely pass through other railroads.
 
Well, surely whom they really use is an "expediter" or "freight forwarder", like the company whose website is shown in the original post (other terms/names are used, too). Railroads are close to natural monopolies - there are only a small number of examples where more than one location is served by more than one rail line - and so to get from continental Point A to B naturally can necessitate using Railroad A, B & C. The expediter does the organization of making the appropriate contracts; when the transportation event occurs, these days normally the entire railcar remains intact and simply is pulled by the different railroad's locomotives across the various sets of lines. That is why you'll see, for example, a Burlington Northern/SP car being pulled by a Chesapeake train, and so forth.
 
Tesla's factory is connected only to Union Pacific. The Gigafactory is also accessible only by Union Pacific. Tesla is, therefore, using Union Pacific. So far it sounds like they're just shipping by rail to Chicago and trucking east of there, so they're probably not using any other railroad. If they decide to ship by rail beyond Chicago, they would have to make arrangements with two railroads, or with a freight forwarder.